The crossmember has the part No C01Q-5019, but this is the assembly which includes various small parts rivetted on. The actual stamping of the centre crossmember is the same as C01T-5019 for the 1940 3 ton truck (and may be the same for smaller trucks eg 1/2 ton, 3/4 ton, 1 ton etc), which would have used these holes to mount the crank handle guide, as surmised by Mike. Will try to find a pic of a commercial truck application.
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